Gray Comma (Polygonia progne)
Gray Comma Polygonia progne Iowa
Gray Comma Polygonia progne Iowa
Gray Comma butterfly (Polygonia progne) on a leaf, dorsal (top), ventral (bottom), at Geode State Park in Iowa.

Details:  Canon EOS 10D,  shutter 1/180 second, aperture f13, ISO 400, Canon EF 100 mm macro lens, Canon MR-14EX flash.
 

Selected References:

Brock Jim P., Kaufman Kenn. Butterflies of North America. New York: Houghton Mifflin Company, 2003. P 198-199.

Struttmann Jane M. USGS Northern Prairie Wildlife Research Center. "Gray Comma  (Polygonia progne)." Butterflies of North America / Butterflies of Iowa. <http://www.npwrc.usgs.gov/resource/distr/lepid/bflyusa/ia/212.htm > (15 Sep. 2004).